Graham received his B.Sc. in Computer Science from the University of British Columbia and subsequently worked on the Ensembl Genome Browser at the European Bioinformatics Institute. Graham then did his Ph.D. with Phil Green at the University of Washington where he researched how selection shapes human genetic variation. During his postdoc with Jonathan Pritchard at the University of Chicago and Stanford, Graham studied how genetic variation affects gene regulation and chromatin. Since January 2016, Graham has been an Assistant Professor at the Salk Institute for Biological Studies.
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